If you have an O/U chambered for 3" shells, you can fire either 2-3/4" or 3" shells through it.
Most field guns are chambered for 3" shells, allowing you to choose which you want to fire, but trap and skeet guns are typically chambered for 2-3/4" shells. Chamber length is usually marked on the barrel.
If the O/U is chambered for 2-3/4" shells, you can only fire 2-3/4" shells through it.
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