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If the primer was indeed positively seated at some point, which you indicate it was, something had to cause it to unseat. Not too many possibilities for that.
You said that the primer fell out on its own. As you also said, it could be an oversized primer pocket, or and undersized primer. It would be easy enough to check the primer pocket on that piece of brass with a micrometer, or by trying to seat a spent primer in it. Positively identifying what happened is your best method of recognizing it if it happens again.
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