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In short, if the state says it is okay - handgun, rifle, or shotgun, concealed or openly carried - it is also okay in National Parks and Wildlife Refuges.
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There's my answer, LOC permitted, if permitted by by state law, right? The way the media has exaggerated the impact of this rule change (though now actually a law), it probably won't surprise anyone much to see Aunt Bessie OCing her .38. After all, they are expecting AKs and ARs, thanks to the alarmists.