It is currently illegal for you to carry a firearm that is readily accessible or loaded in a National Park. The only way to legally possess a firearm in a National Park is if it is inaccessible (locked up) or broken down to a non-functioning state; AND unloaded. Permit status has nothing to do with it.
That will change on February 22, 2010. On or after February 22, 2010 you may carry your firearm in the National Park according to any state law that the National Park is located in. If you are in the Idaho section of Yellowstone, you carry according to Idaho laws, if you are in the Wyoming section of Yellowstone, you carry according to Wyoming. In states that allow carry without a permit, you may carry in the National Park without a permit as well on or after February 22, 2010.
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