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The length of the cone can be important. The shorter cone will mean the hammer face will not strike the cap properly; the hammer face is supposed to be parallel with the top of the cone, and thus strike the cap evenly all around. The shorter cone means that only a part of hammer face will strike the cap. In an extreme case this could mean FTF's. It could also lead to hot gas blowing back through the nipple flash hole and escaping into the hammer area since part of it won't hold the cap down. It might even mean the hammer face doesn't contact the cap at all - the alignment might be so bad that the lip of the hammer hits the cone preventing the face from reaching the nipple.
The real dimension to check is not the length of the cone but the distance from the surface where the nipple seats against the gun to the top of the cone. The flange on the shorter cone nipple might be thicker and make up for the shorter cone.
1/4x28 threaded nipples come in several lengths; the shorter ones are generally for pistols, with the longer ones usually used on rifles. That's not always true, however. Sounds like your Howdah uses standard rifle nipples.
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