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I'm not even sure it's the polygonal rifling that causes the problem. It very well could be a sharper shoulder at the front of the chamber & case mouth that allows the build up of lead particles/shavings during firing, and then starts to push each next round further back, until the cartridge fires from a less supported breech.
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Ding! Ding! Ding! Ladies and gentleman we have a winner.
And for those of you who wonder about polygonal rifling guess what, it was around long long before jacketed bullets.