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That serial number very likely means that it's a .455 Mark II Second Model, manufactured between 1915-1917. The majority of these guns went to the British, but over 15,000 went to Canada.
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I have one of these, and it is not marked .455 Colt, but merely .455. Mine has been converted to .45 ACP, and the second "5" in .455 has been X-d out and an "AR" (Auto Rim) stamp added. I have WWII-era Canadian military ammo, and it does fit in my Webley Mk. I, which makes sense, since the later cartridge has a shorter case. My S&W has a M1917 cylinder fitted, so I can't make any determination about the original chambering.